Falconer's formula Created 2026-09-28 Updated 2026-11-05
Falconer's formula estimates narrow-sense heritability from monozygotic and dizygotic twin correlations:
It relies on equal relevant environments and a dizygotic additive-genetic correlation of one half.
The common marginal trait variance is
For monozygotic pairs, shared genes and shared environment give
For dizygotic pairs, the genetic covariance is halved while the common environment is unchanged:
Subtracting cancels the common-environment variance and proves Falconer's formula
the heritability under the ACE model.