= J. B. S. Haldane
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<Geneticist> who analyzed the relationship between <sex chromosomes> and <hybrid sterility>. Haldane’s 1922 paper identified the recurring association of absent, rare or sterile <hybrid> offspring with the <heterogametic sex>.<Sex ratio and unisexual sterility in hybrid animals by J. B. S. Haldane>[Title, author list and reported study]{ref}
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{title=Portrait of <J. B. S. Haldane>}
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