= Jonathon Howard
{c}
A researcher in <biophysics> who studies how <kinesin> and other molecular machines generate forces and organize the <cytoskeleton>. Measurements on individual molecules help connect their mechanical action with cell movement and shape.<Professional profile of Jonathon Howard at Yale University>[Biographical Sketch: motor proteins, cytoskeleton and single-molecule measurement techniques]{ref}
He also coauthored a 2006 <optical tweezers> calibration study, providing quantitative position and force scales for such measurements.<Calibration of optical tweezers with positional detection in the back-focal-plane by Simon F. Tolić-Nørrelykke and others>[Author list and abstract: force and position calibration]{ref}
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