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Yes. An immobile $Y$ concentration can still change locally through reaction while diffusing $X$ transports activation. Vanishing diffusion of one component does not prohibit a propagating reaction front.

In the invariant special case $v=0$, the first equation is already Fisher's equation with growth coefficient one, so it has travelling fronts regardless of $D$. A one-diffuser reaction front also gives a coupled example with nonzero $v$. For $z=\xi -c\tau$, the equations with $D=0$ areFor $0<\delta <2$, choose $k=\sqrt{1-\delta /2}$, $\beta v_0=2-\delta$, and setIndeed $U^{\prime }=kU(1-U)$ and $V^{\prime }/V=-2kU$, giving the second equation. In the first, division by $U(1-U)$ leaves $k^2(1-2U)+ck+1-\beta v_0(1-U)$, whose constant and $U$ coefficients both vanish with these choices. Ahead of the left-moving wave $U=0,V=v_0$; behind it $U=1,V=0$. This explicitly demonstrates a travelling front with substrate depletion but no $Y$ diffusion, rather than relying solely on the uncoupled special case.