Solution
= Solution
Of the $a_t$ immature cells, a fraction $\lambda$ leaves by maturation and the rest remains immature. Of the $b_t$ mature cells, a fraction $\mu$ divides and is replaced by four immature cells, while a fraction $k$ dies; the remainder stays mature. These contributions give exactly
$$
\boxed{a_{t+1}=(1-\lambda)a_t+4\mu b_t,
\qquad
b_{t+1}=\lambda a_t+(1-\mu-k)b_t.}
$$