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Write proper position as $\mathbf{r}=a(t)\mathbf{x}$ and proper velocity asSubstitute this decomposition into the proper-coordinate Euler equations for an inviscid fluid, use ${\nabla }_r=a^{-1}{\nabla }_x$, and subtract the homogeneous-background acceleration. With $\rho =\bar{\rho }(1+\delta )$, one obtains the comoving peculiar-velocity equationThe peculiar gravitational potential $\Phi$ is the total Newtonian potential with the potential of the exactly homogeneous expanding background subtracted. Its gradient therefore generates only accelerations relative to the Hubble flow.

For a pressureless fluid, linearization discards the quadratic advection term, leavingIn the early Einstein-de Sitter universe, the growing mode has $\Phi (\mathbf{x},t)=[D(a)/a]{\Phi }_i(\mathbf{x})$. Integration gives

The linear growth factor obeysSince $\bar{\rho }={\bar{\rho }}_i/a^3$ after choosing $a_i=1$, this is equivalent toIt follows that $a\mathbf{v}=-a^2\dot{D}\nabla {\Phi }_i/(4\pi G{\bar{\rho }}_i)$. Since $\dot{\mathbf{x}}=\mathbf{v}/a$, one final integration gives the Zel'dovich approximationwhere an additive initial displacement has been absorbed into ${\mathbf{x}}_i$.