= SHATTERPROOF2
{c}
<SHATTERPROOF2> is an <Arabidopsis thaliana> <gene> encoding a <MADS-box transcription factor> that acts with <SHATTERPROOF1> in <fruit dehiscence>. The 2000 study connected their combined activity to <dehiscence zone> differentiation and <lignin> deposition in adjacent <cells>.<SHATTERPROOF MADS-box genes control seed dispersal in Arabidopsis by Sarah J. Liljegren and others>[Abstract; mutant and constitutive-expression studies]{ref}
Loss-of-function and constitutive-expression experiments ask complementary questions: what fails when the normal regulators are absent, and what changes when their activity is supplied outside its usual pattern? Neither question identifies the <transcription factor>'s direct <DNA> targets by itself. Establishing regulation of tissue development is a different evidential step from identifying each directly bound <gene>.
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