= Species concept
A <species concept> states what should make <organisms> members of the same <species>; an operational criterion states what observations will test that claim. <Reproductive isolation>, distinguishable <morphology> and ancestry inferred through <phylogenetics> answer different questions. In a lineage-based interpretation, they are evidence for separation of evolving <biological populations>, and need not appear together. <Species Concepts and Species Delimitation by Kevin de Queiroz>[pp. 880–883, Table 1, Figure 1 and “Conceptualization versus Delimitation”: distinguish defining properties from operational evidence]{ref}
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