Nature458(7237),475–480(2009), DOI10.1038/nature07851; PMID19325628. Primary first-author byline Daniel A.Pomeranz Krummel verified. Abstract, Structure determination and Figures1–4: reconstituted U1 functional core, seven Sm proteins, heavy-atom labeling and 5.5angstrom map. Engineered/truncated particle lacks U1A; Figure5 adds the known U1A–RNA structure to model a complete particle. The crystallographic core must not be described as every component of an intact native U1 particle.