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Use the mostly-positive target Minkowski spacetime metric and a closed string coordinate $0\le \sigma <2\pi$. In conformal gauge, variation of the Polyakov action gives $({\partial }_{\tau }^2-{\partial }_{\sigma }^2)X^{\mu }=0$. The compact coordinate, denoted $X^{25}$, can change by $2\pi Rw$ around the closed string, with $w\in \mathbb{Z}$. Write $w^{\mu }=w{\delta }^{\mu ,25}$. Integrating the two chiral wave solutions and expanding their periodic parts gives the closed-string mode expansionReality imposes ${\alpha }_{-n}=({\alpha }_n{)}^{\ast }$ and the analogous condition in the other chirality. The classical compact momentum is continuous; quantization of a wavefunction on the circle subsequently gives $p^{25}=n/R$ with $n\in \mathbb{Z}$. The momentum and winding modes are most conveniently packaged asFor noncompact directions both zero modes are $\sqrt{{\alpha }^{\prime }/2}{p}^{\mu }$. Thus the formula includes both independent string oscillator families and all compact topological sectors.

Define the worldsheet stress-energy tensor by ${\theta }_{ab}=-2(-\gamma {)}^{-1/2}\delta I/\delta {\gamma }^{ab}$. Its normalization and components areAn overall rescaling of the named worldsheet stress tensor does not alter its vanishing. Variation of the independent worldsheet metric imposes ${\theta }_{ab}=0$, equivalently $(\dot{X}+X^{\prime }{)}^2=(\dot{X}-X^{\prime }{)}^2=0$. These are the Virasoro constraints that restrict the otherwise general wave solution.

The conjugate momentum density is $P_{\mu }={\dot{X}}_{\mu }/(2\pi {\alpha }^{\prime })$. The gauge-fixed Hamiltonian generating $\tau$ translation is thereforewith the analogous tilded generators. In particular, before imposing the Virasoro constraints,The target momentum includes its timelike component, so this is not a positive particle-energy formula. On the classical constraint surface the reparameterization Hamiltonian vanishes; target energy is the separate charge $p^0$. The difference $L_0-{\tilde{L}}_0=0$ imposes the residual spatial constraint.

In the quantum theory the same quadratic expressions require normal ordering: $[{\alpha }_m^{\mu },{\alpha }_n^{\nu }]=m{\eta }^{\mu \nu }{\delta }_{m+n,0}$ makes different orderings differ by divergent constants. Their regularized finite remainder is the normal-ordering constant of a string, or string intercept. The Virasoro central extension also appears. In the critical 26-dimensional bosonic string theory, the 24 transverse string oscillators give one-unit string intercept in each chirality. For example their regulated chiral vacuum energy is $24{\sum }_{k>0}k/2=-1$ using zeta function regularization. The physical zero-mode constraints are $(L_0-1)|\Psi ⟩=({\tilde{L}}_0-1)|\Psi ⟩=0$.

Let $M$ be the mass seen in the remaining 25 noncompact dimensions. Substituting the compact zero modes in these two constraints givesTaking their mean and difference yieldsThis sign of compact-circle closed-string level matching follows from assigning tildes to the left-moving sector. Under T-duality, set $R^{\prime }={\alpha }^{\prime }/R$, $n^{\prime }=w$, $w^{\prime }=n$. Then $p_L^{\prime }=p_L$ and $p_R^{\prime }=-p_R$, so reversing the compact right-moving string oscillators implements the transformation while leaving $N,\tilde{N}$ unchanged. Both the mass equation and closed-string level matching are invariant. This proves invariance of the full closed string spectrum, including its string oscillator degeneracies, rather than only the zero-mode contribution.

A D-brane with $p$ spatial worldvolume directions, called a D$p$-brane, is a $(p+1)$-dimensional timelike submanifold on which open string endpoints have Neumann boundary conditions tangentially and Dirichlet boundary conditions transversely. Apply T-duality to the compact coordinate by $\tilde{X}=X_L-X_R$. The differential relationsinterchange the two endpoint conditions. Consequently a D$p$-brane wrapping the original circle becomes a D$(p-1)$-brane localized on the dual circle. Its compact Wilson line specifies that position. This is T-duality of D-brane boundary conditions.

The separated-brane configuration must instead have the circle transverse to the D-branes. Two parallel D-branes wrapping the same circle cannot be separated along that worldvolume direction. Thus the latter part describes a new orientation, with both endpoints subject to compact Dirichlet boundary conditions. If their positions differ by $L$, an oriented Open string stretched between D-branes has displacementOn $0\le \sigma \le \pi$, its compact classical solution has a linear part $y_1+L_w\sigma /\pi$, plus sine modes vanishing at the endpoints. This adds $L_w^2/(4{\pi }^2{\alpha }^{\prime })$ to the Virasoro generator zero mode. Thus, for momentum $\mathbf{p}$ along the D-branes,The second equation is the quantum rest-energy squared of the string oscillator ground state; the smallest one uses the shortest allowed compact separation $|L_w|$. Classically the straight string energy is $T|L_w|=|L_w|/(2\pi {\alpha }^{\prime })$. The quantum ground state is a tachyon if $|L_w|<2\pi \sqrt{{\alpha }^{\prime }}$, so in that regime it is an instability rather than a state with real rest energy. Dropping the string intercept would describe the classical stretched string, not its quantum ground state.

The transverse-circle T-duality changes these two D-branes to D$(p+1)$-branes wrapping the dual circle. Their positions become the eigenvalues $A_i=y_i/(2\pi {\alpha }^{\prime })$ of a constant gauge connection. The Wilson loop phases are $e^{i2\pi R^{\prime }{A}_i}=e^{i{y}_i/R}$. A string connecting D-brane $1$ to D-brane $2$ has dual compact momentumIts momentum contribution to $E^2$ is exactly the former stretching term, including every winding sector. Qualitatively the two geometrically coincident wrapped D-branes are distinguished by their Wilson lines, and their connecting strings have shifted momentum quantization. This is how brane separation becomes a Wilson line.