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Coauthor of research clustering predicted protein structures at very large scale. The work connected structural similarity with organization of the known protein universe, complementing comparisons based only on sequence.^[ref]

Using the supplied cluster counts,These are structural clusters represented across the three domains of life under the stated classification. They are not necessarily folds present in every individual species, nor must every cluster be a distinct topological fold. The distinction between a cluster and a fold depends on the structural-similarity criterion. The underlying large-scale analysis explicitly used structural clustering of predicted models.^[ref]