TODO: biological interpretations and organism-specific propagation are user-deferred. The notes below concern only generic excitable reaction–diffusion systems.
A front solution joins different asymptotic states in space. In a reaction–diffusion system, local nonlinear dynamics select possible states while diffusion couples neighboring points. A travelling wave with profile , , converts a partial differential equation into a boundary value problem for the shape and speed. A pulse instead returns to the same resting state on both sides and can contain leading and trailing fronts.
A useful nondimensional cubic fast-slow reaction-diffusion model, in a convention of the FitzHugh-Nagumo model, iswhere , , and . Here is a fast activation variable, a slow negative-feedback variable and a generic drive; no physiological variable is assigned. Without spatial coupling, nullclines are and . At , the resting state has Jacobian matrixIt has linear stability. With appropriate parameter choices a sufficiently large perturbation makes a fast excursion to the other outer branch of the cubic, followed by slow feedback and return. Thus a stable rest state can be excitable without being linearly unstable. Other parameters or applied drive can yield multiple equilibria or oscillations; these behaviours should not be asserted for every parameter set.
The fast-subsystem front can be obtained explicitly by freezing . For the bistable cubic reaction-diffusion equation, are stable and is unstable. The decreasing logistic front of a bistable cubic equation isTo check it, and substitution into cancels every term. For , the high state advances to the right; for , the direction reverses. At the front is stationary. More generally, multiplying the travelling-wave equation by and integrating yieldsThis links propagation direction to the bias between the two stable states and recovers Maxwell balance for a scalar reaction-diffusion front at zero speed. It is a bistable selected-speed problem, rather than a universal Fisher-type minimum-speed formula.
For the coupled fast-slow system, travelling profiles satisfyA slow recovery wake can support a return front behind the advancing fast front, producing a travelling pulse for suitable parameters. Its width reflects the slow recovery time and propagation speed, while each sharp edge has the faster diffusion–reaction scale. The frozen-recovery speed above is a controlled subsystem result, not an exact speed for every coupled pulse. A definite pulse speed requires parameters and connecting boundary condition; none are specified by this brief-note question.
