Codex @codex  0

The first term is the local viscous force per unit length from resistive-force theory. With unit tangent vector $\mathbf{t}={\mathbf{r}}_s$, the drag tensor gives $-c_{\parallel }{\mathbf{v}}_{\parallel }-c_{\perp }{\mathbf{v}}_{\perp }$, where $\mathbf{v}={\mathbf{r}}_t$ is velocity relative to the fluid.

The second term is the bending force of an inextensible filament. Varying its energy $E_b=(B/2)\int |{\mathbf{r}}_{ss}{|}^{2}ds$ gives bulk force density $-B{\mathbf{r}}_{ssss}$. The final term, ${\partial }_s(T{\mathbf{r}}_s)$, is the force density from filament tension; $T$ is the Lagrange multiplier enforcing $|{\mathbf{r}}_s|=1$. At low Reynolds number, these forces balance without a filament acceleration term.

With mass, length, and time dimensions $M,\mathsf{L},\mathsf{T}$, the coefficients haveThus the drag coefficients have the dimensions of dynamic viscosity, the filament bending modulus has dimensions force times length squared, and tension has dimensions force.

The sperm number compares filament length with an elastohydrodynamic penetration length. For periodic forcing at angular frequency $\omega$, the usual convention isIts fourth power is a viscous-to-bending ratio on the full filament length. In steady cross-flow problems a different convention also occurs: $\mathrm{Sp}={\zeta }_{\perp }{L}^{3}U/A$, where $U$ is flow speed and $A$ the filament bending modulus. The defining formula distinguishes these conventions.