= Joshua Lederberg
{c}
{title2=1925–2008}
<Geneticist> who, with <Esther Lederberg>, developed <replica plating> to investigate <bacterial> <mutants>. Preserving the spatial pattern of colonies allowed selection to reveal variants that already existed before exposure.<Replica plating and indirect selection of bacterial mutants by Joshua Lederberg and Esther Lederberg>[Title, author list and reported study]{ref}
\Image[https://upload.wikimedia.org/wikipedia/commons/a/af/Joshua_Lederberg.jpg]
{title=Portrait of Joshua Lederberg}
{width=220}
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