In the dark, energy minimization gives . Translation and rotation are zero-energy freedoms; one representative is . After illumination, choose the equivalent light-adapted equilibriumThen has homogeneous free-end conditions , and its initial valueis orthogonal to the two rigid zero modes and .
Let solve the free--free biharmonic eigenvalue equationand chooseThese are orthogonal eigenfunctions of the biharmonic operator with at both ends. The shape iswhereThe omitted zero modes would only translate or rotate the whole filament.
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