For each Volvox colony, the excess gravitational force isFor a normal Stokeslet a distance below a flat stress-free interface, the image is an oppositely directed Stokeslet a distance above it. The real force creates no lateral velocity at the other colony because both centres have the same height. Their separation from the image is , however, so each colony moves toward the other with speedBoth colonies move, and therefore
DefineThe dimensionless dynamics areThis gradient flow descends an even attractive potential with minimum and as .
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