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The quaternionic projective space ${\mathbb{HP}}^n$ is the space of one-dimensional right quaternion subspaces of ${\mathbb{H}}^{n+1}$. Equivalently it is the quotient of the unit sphere $S^{4n+3}$ by simultaneous right multiplication by unit quaternions. Its coordinate filtration has one open cell ${\mathbb{H}}^k\cong {\mathbb{R}}^{4k}$ in each dimension $4k$, for $0\le k\le n$. Hence its cellular cohomology is $\mathbb{Z}$ in those dimensions and zero otherwise.

Let $\mathcal{H}$ be the quaternionic tautological line bundle. Its unit sphere bundle is $S^{4n+3}\to {\mathbb{HP}}^n$, with fibre $S^3$. The Gysin sequence of a sphere bundle shows that multiplication by its Euler class $e\in H^4$ is an isomorphism from $H^{4k}$ to $H^{4k+4}$ for $k<n$. Choose the generator $u=-e$. Its powers generate every nonzero positive degree, giving the cohomology ring of quaternionic projective spaceThis also accounts for $n=0$.

First take $n=2$. Under the coordinate inclusion $i:{\mathbb{CP}}^2\to {\mathbb{HP}}^2$, the pulled-back quaternionic line is the quaternionic extension of the complex tautological line $L$. As a complex rank-two bundle it is $L\oplus \overline{L}$: a transition scalar $c\in \mathbb{C}$ acts on the two complex coordinates of a quaternion by $c$ and $\overline{c}$. Put $x=c_1(L^{\ast })$, the degree-two generator of the cohomology ring of complex projective space. The Whitney sum formula for Chern classes givesHere the Euler class of a complex vector bundle is its top Chern class, using the complex orientation. In particular this degree-four pullback has coefficient one; it is not a multiple of larger absolute value. The compatible tautological bundles on the projective filtrations give the same equality for every $n$, and multiplicativity then determines the whole ring map:It is zero whenever $2k>n$. These facts are the complex inclusion into quaternionic projective space.

For an odd prime $p$, the Steenrod reduced powers are natural stable cohomology operationsThey satisfy $P^0=1$, the Cartan formula, $P^{i}a=0$ when $2i>|a|$, and $P^{i}a=a^p$ when $|a|=2i$. In particular, on ${\mathbb{CP}}^{\infty }$ one has $P^{1}x=x^p$ and $P^{i}x=0$ for $i>1$. The Cartan formula and the binomial theorem givePass to the infinite projective spaces, where $i^{\ast }:H^{\ast }({\mathbb{HP}}^{\infty };{\mathbb{F}}_p)\to H^{\ast }({\mathbb{CP}}^{\infty };{\mathbb{F}}_p)$, $u\mapsto x^2$, is injective. The equality just obtained determines the Steenrod powers on quaternionic projective space; restricting to the finite spaces giveswith coefficients modulo $p$ and powers above $n$ set to zero. In particular $P^{1}u=2u^{(p+1)/2}$, $P^{2}u=u^p$, and $P^{i}u=0$ for $i>2$. The infinite-space argument matters: the finite inclusion cannot detect those degrees for which $x^{2k}=0$.

Finally put $Q={\mathbb{HP}}^{15}/{\mathbb{HP}}^{12}$ and $S={\Sigma }^{48}{\mathbb{HP}}^3$. Both have reduced cohomology $\mathbb{Z}$ in degrees $52,56,60$ and zero otherwise. Choose integral generators $a_{13},a_{14},a_{15}$ for $Q$ whose pullbacks under the quotient map are $u^{13},u^{14},u^{15}$, and suspended integral generators $b_1,b_2,b_3$ for $S$ from $u,u^2,u^3$.

Use $p=5$. Naturality for the quotient and the formula above giveStability under the suspension isomorphism instead givesAny homotopy equivalence $h:Q\to S$ would induce isomorphisms on the rank-one integral groups, so $h^{\ast }{b}_1=\epsilon a_{13}$ and $h^{\ast }{b}_3=\delta a_{15}$ with $\epsilon ,\delta \in \{1,-1\}$. Reducing modulo five and commuting $h^{\ast }$ with $P^1$ would require $\epsilon =2\delta$ in ${\mathbb{F}}_5$. Neither $1$ nor $-1$ equals $2$ or $-2$ modulo five. ThereforeThe essential point is that integral generator signs constrain Steenrod comparisons. Arbitrary changes of basis over ${\mathbb{F}}_5$ could rescale these two nonzero coefficients into agreement; a genuine equivalence must also preserve the integral lattices, where only the two signs are available.

Its fibre over a point of quaternionic projective space is the quaternionic line represented by that point. As a real vector bundle it has rank four, and its unit sphere bundle is $S^{4n+3}\to {\mathbb{HP}}^n$. Right multiplication by a chosen imaginary unit equips it with complex rank two. Under the complex inclusion into quaternionic projective space, it restricts to $L\oplus \overline{L}$, where $L$ is the complex tautological bundle. Thus its Euler class restricts to $-c_1(L^{\ast }{)}^2$.

Choose the degree-four generator with pullback $u\mapsto x^2$ under the complex inclusion into quaternionic projective space. On infinite complex projective space, the Cartan formula gives $P^i(x^{2k})=\left(\genfrac{}{}{0ex}{}{2k}{i}\right)x^{2k+i(p-1)}$. Injectivity of the infinite-space pullback proves the displayed formula for quaternionic projective space. Restriction to ${\mathbb{HP}}^n$ sets powers above $n$ to zero. Coefficients are reduced modulo the odd prime $p$; the exponent is an integer because $p-1$ is even.