= Temporal sensing in bacterial chemotaxis
<Temporal sensing in bacterial chemotaxis> compares changing stimulation during movement with a recent adapted baseline. Attractant increases can lengthen runs and decreases can increase tumbling, producing biased movement up a chemical <gradient>. Rapid spatially uniform concentration steps distinguish a temporal response from a requirement for front-to-back spatial comparison.<The gradient-sensing mechanism in bacterial chemotaxis by Robert M. Macnab and Daniel E. Koshland Jr.>[pp. 2509–2512, temporal gradient apparatus and motility responses]{ref}
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