Temporal sensing in bacterial chemotaxis compares changing stimulation during movement with a recent adapted baseline. Attractant increases can lengthen runs and decreases can increase tumbling, producing biased movement up a chemical gradient. Rapid spatially uniform concentration steps distinguish a temporal response from a requirement for front-to-back spatial comparison.[ref]
The gradient-sensing mechanism in bacterial chemotaxis by Robert M. Macnab and Daniel E. Koshland Jr.. pp. 2509–2512, temporal gradient apparatus and motility responses
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