 Bipes

Bipes is a taxonomic genus of burrowing amphisbaenians distinguished by retaining external forelimbs despite an elongated body and absence of external hindlimbs. It is an informative comparison with other amphisbaenians, which lack external limbs: external appearance alone does not reveal how much internal anatomy remains.^[ref]

In a 2019 anatomical study, investigators used iodine-enhanced computed tomography to distinguish soft tissues and reconstruct shoulder muscles as well as bones. Bipes retained an almost complete shoulder girdle, while other sampled amphisbaenians had much more reduced bony structures. Nevertheless, all sampled amphisbaenian forms retained the full set of 17 shoulder muscles, sometimes fused or rearranged. Muscle attachment could shift from bone to surrounding soft tissue.^[ref]

The result challenges a tempting shortcut: losing a visible limb does not necessarily delete all its associated muscles at once. Bipes supplies the limbed comparison that helps reveal which internal components have persisted or changed during the evolution of the burrowing body form.