A Miocene fossil ape with a mosaic of trunk and hand characters. Its informative anatomy does not uniquely identify it as the Pan–Homo last common ancestor. [ref]
Pierolapithecus catalaunicus, a new Middle Miocene great ape from Spain by Salvador Moyà-Solà and others. Science 306, pp. 1339–1344; orthograde trunk and contrasting hand morphology
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