A type species is the species serving as the nomenclatural type of a genus under a naming code that uses this form of typification. It anchors application of the genus name. It is not necessarily an average, an ancestor or the most familiar member of the genus.
In prokaryotic nomenclature, the Code explicitly identifies the type of a genus or subgenus with a type species and specifies how that species is fixed. If a genus originally included only one species when validly published, that species is its type even without an explicit designation. These are naming rules, not measurements of evolutionary distance.[ref]
International Code of Nomenclature of Prokaryotes. Prokaryotic Code (2025 Revision) by Aharon Oren and others. Rule15 and Rules20a–20c, printed pp.10 and13: types, designation and a genus originally containing one species
An original hypothetical example separates the reference from a summary statistic. Suppose a genus contains species T, U and V, with T the designated type species. A measured trait has values1,10 and11, whose arithmetic mean is . U lies nearer that mean, but this does not make U the type. The designation and the statistical comparison answer different questions.
Now suppose new evidence from phylogenetics divides the hypothetical group into T+U and V. The existing reference is associated with the group containing T; how the remaining group is named requires the relevant naming rules and publication history. Merely discovering that V has an unusual trait does not transfer the reference to it.
The anchor has another level beneath it. In the ordinary prokaryotic case a species itself has a type strain, whereas the genus has a type species. A stored culture accession, a species reference and a genus reference therefore do not all denote the same rank or the same physical object.[ref]
International Code of Nomenclature of Prokaryotes. Prokaryotic Code (2025 Revision) by Aharon Oren and others. Table2 under Rule15 and Rule18a: species and genus types; Rule20a: type species
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