Let be prey and predator abundances. With constant positive demographic and predation coefficients, the Lotka-Volterra equations are
Their positive equilibrium is , . Its Jacobian matrix and eigenvalues are
The linearized populations oscillate with period , with the predator peak following the prey peak. This is a center equilibrium, not an attracting equilibrium. The conclusion is supported beyond linearization by the Logarithmic first integral of the Lotka-Volterra equations: in these units one may use
Direct differentiation gives . Its Hessian matrix is a positive-definite matrix in the positive quadrant, and its minimum is at . Nearby level curves are closed periodic orbits, so there is no damping toward a preferred oscillation amplitude.
For , take the Logarithmic first integral of the Lotka-Volterra equations
Along a trajectory,
Thus is a first integral. The scalar function is nonnegative for , vanishes only at , and tends to infinity as or . Moreover,
is positive definite, so is strictly convex. It follows that is its unique critical point and every level set is a compact smooth simple closed curve in the positive quadrant. The vector field is nonzero on such a level curve, so uniqueness of solutions makes each connected level curve one periodic orbit. The zero level consists of the fixed point . Hence every trajectory in is either periodic or the fixed point .