Use the rotation convention of part (a), so the orientation drift is . The steady gyrotactic orientation Fokker-Planck equation on the unit sphere isLet , , and introduce the rotation generator . Dividing by givesThe leading isotropic density is . At first order, writing , rotational diffusion makes no contribution because . The tangential drift is the spherical gradient of , whose divergence is . ThereforeIn the polar convention of the paper, , and . Thus, in the specified first-harmonic form,The correction integrates to zero, preserving normalization. The first correction has no dependence on because the isotropic leading density has zero spherical Laplacian; enters the next-order equations.
Spherical symmetry gives . Consequently the rapid-rotation mean gyrotactic orientation isIt is this small horizontal swimming bias, rather than a mean vertical orientation at leading order, that enters the concentration equation of part (c).
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