A fixed comoving volume labels a fixed collection of cold dark matter particles. Their background mass is , which is constant by mass conservation. At the perturbed positions it is . The Jacobian determinant of the Zeldovich approximation map isTaking its reciprocal and using mass conservation givesFor the perturbed field the Poisson equation must be . The displayed background-only right-hand side in the PDF would give no clumping force; it is the background equation, not the full perturbed equation.
Write , where . To linear order,For the longitudinal, curl-free displacement used in the growing Zel'dovich mode, this integrates to , after removing external homogeneous harmonic fields. More generally the force involves only the longitudinal part , which has the same divergence as .
Differentiate twice at fixed and substitute . The terms proportional to cancel using , leavingFor a longitudinal displacement the right side is . Taking minus the comoving divergence gives the general scalar density contrast equation
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