A seed dispersal syndrome is a combination of seed or fruit traits associated with a mode of seed dispersal. Wings can favor transport by wind, while fleshy fruits can attract animals that later deposit the enclosed seeds. The traits generate a hypothesis about likely transport, not direct proof of the agent responsible for every dispersal event.
A comparative study connecting fruit traits with observed disperser networks found that morphological traits better supported broad groups of dispersing birds, terrestrial mammals and bats than chemical or display traits did.[ref]
Fleshy fruit traits and seed dispersers: which traits define syndromes? by Tobias Nicolas Rojas and others. Annals of Botany 129, pp. 831–838; trait groups and disperser functional types
Direct observations remain valuable because removal of a fruit, deposition of a viable seed and establishment after germination are different stages of successful seed dispersal. Articles by others on the same topic
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