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A microtubule-organizing structure in relevant animal cells. Positioning centrosomes relative to a niche can prepare the orientation of a later mitotic spindle. Budding yeast instead has a spindle pole body.

Its microtubule-nucleating material helps organize arrays without requiring every filament to remain permanently attached. A centrosome and a centriole are different structural levels: centrioles can be components of a centrosome, while the surrounding material carries much of the organizing activity. ^[ref]