Seed germination is renewed plant embryo activity and growth. Many germination assays score completion as radicle emergence, before seedling establishment. Seed imbibition precedes that endpoint; swelling alone is insufficient.[ref]
Seed dormancy and the control of germination by William E. Finch-Savage and Gerhard Leubner-Metzger. Seed dormancy and the control of germination, §II.1, p. 502: water uptake, embryo expansion and radicle protrusion
Water, molecular oxygen and suitable temperature support germination; light sensitivity depends on the species and seed state. Neither illumination nor one particular temperature is universally required. Seed dormancy connects germination timing to ecology by delaying seedling establishment.[ref]
Seed dormancy and the control of germination by William E. Finch-Savage and Gerhard Leubner-Metzger. Seed dormancy and the control of germination, Summary and §II.1–2, pp. 501–502: environmental requirements and timing
The distinction matters when comparing propagation experiments: a treatment that increases radicle emergence need not improve later seedling survival. Define the endpoint before comparing treatments, then measure the later outcome separately.
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