Solution

ID: past-exam-of-the-mathematics-and-the-environment-and-mathematical-physiology-course-of-the-university-of-oxford/2009/a/1/v/solution

For , the positive equilibrium is
Set and and retain first-order terms. The linear system is
Differentiating its first row and using the second gives
In particular the trace is positive and determinant is . The steady drainage state is linearly unstable for every positive supply, whether its instability is oscillatory or nonoscillatory.

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