A force-free, torque-free microswimmer in a stationary Newtonian fluid, with negligible fluid and body inertia and steady Stokes flow at each instant, gains no net displacement from a reciprocal stroke: one whose sequence of shapes is exactly retraced in reverse. The environment and boundary conditions must also be unchanged during that retracing. Opening a two-shell swimmer and then closing it follows the same sequence backwards; doing the closure faster does not help, because Linearity of Stokes flow changes the instantaneous velocity by the same factor as the stroke rate.
A single real shape coordinate that travels out and back along an interval gives this cancellation, as computed in the force-free two-sphere stroke. Two independently controlled shape coordinates can instead execute a loop that is not retraced, avoiding this obstruction. Continuous rotation also need not retrace an interval: the helical microswimmer with a spherical head provides another route to propulsion. The scallop theorem is a restriction on a stroke under stated conditions, not a claim that small organisms cannot swim. Purcell presented the opening-and-closing scallop and a swimmer with two hinges in Life at low Reynolds number (1977), figures 6–7.
The Scallop Theorem is a concept from the field of mathematical biology, specifically in the study of the dynamics of movement in organisms. It addresses the limitations of locomotion in certain types of organisms, particularly those that are at or near the microscopic scale, like small aquatic animals or microorganisms. The theorem states that certain types of organisms cannot swim effectively by using only passive movements in their appendages, such as flagella or cilia.
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