A family of models in which an instructive signal changes the influence of recently active inputs on cerebellar output. In a smooth-pursuit experiment, complex-spike duration predicted subsequent simple-spike depression and eye-movement learning; matching preceding simple-spike activity provided a control. This supports graded instruction during behaviour without proving an exact delta rule at every synapse.[ref]
Purkinje-cell plasticity and cerebellar motor learning are graded by complex-spike duration by Yan Yang and others. Figures 1–4 and Methods; author manuscript at PMC4132823: smooth pursuit, trial-over-trial plasticity and controls for prior simple-spike activity
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