For a barotropic fluid, the squared sound speed is . A Fourier density mode obeys
in linear Newtonian perturbation theory.
For a barotropic perfect fluid with constant , its cosmological adiabatic sound speed satisfies . If , the short-wavelength pressure term has the wrong sign for restoring oscillations: a mode grows as when expansion can be neglected. For in a coasting background with constant conformal expansion rate , the exponents are . This is a gradient instability, not ordinary positive-pressure Jeans instability. A network with tension or elastic anisotropic stress needs different constitutive perturbation equations; a negative background equation of state alone does not establish its perturbation sound speed.
The squared adiabatic sound speed is the ratio of background pressure and energy-density time derivatives. For a barotropic equation of state it is . The non-adiabatic pressure perturbation is the difference ; in a general field or multicomponent model this background quantity need not equal the rest-frame propagation speed.

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