Let be the density contrast, let be a comoving coordinate, and use . Define , so for the pressureless background. The perturbation Poisson equation isFor the comoving velocity field , local mass conservation givesOne obtains this by subtracting from . In the linear regime it reduces to , the linearized cosmological continuity equation. The spatial derivatives are with respect to , and the time derivative in the conservative continuity equation is at fixed . In contrast the acceleration of an individual fluid element is a material derivative. The physical peculiar velocity is .
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