Peristaltic pumping transports fluid by a travelling deformation or travelling external pressure acting on a tube. With wave coordinate , mass conservation makes the relative flux constant. The laboratory flux is , so a negative relative flux does not rule out forward mean pumping.
For negligible inertia, periodic forcing , resistance and prescribed mean area , write . First-order area obeys . Its mean square is . Averaging the second-order friction balance gives , proving the displayed forward pumping rate. Periodicity entails zero mean pressure drop; the mean-area convention fixes the expansion's otherwise free constant.

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