Q. Gu is the first-listed author of the 1998 study identifying the developmental role of FRUITFULL in Arabidopsis thaliana. The paper connected a disrupted MADS-box transcription factor gene with defective cell differentiation in fruit valves. Its indexed byline gives Q. Gu; an expanded given name is not established here.[ref]
The experiment made the mutation informative in two ways. An enhancer trap insertion disrupted the gene, while its reporter gene marked expression in valve tissues. In the mutant, valve cells failed to elongate and differentiate normally; the mature fruit consequently remained short and its tissues grew without normal coordination.[ref]
The combination is more revealing than organ length alone: a local expression pattern and a tissue-specific defect connect a regulatory gene to development at the cell level. They support a role in the process without demonstrating that every downstream growth gene is a direct transcriptional target.

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