In the travelling coordinate , mass conservation becomes . Thus the conserved relative flux is and
For a periodic peristaltic pumping solution, the neglected-inertia momentum equation is
Take to be the spatial mean area, so . This normalization, and zero net pressure drop over a period, are implicit in the specified periodic forcing. The constant leading state has no pressure gradient, giving . Put and . At first order,
Period averaging gives , and the zero-mean solution is
For the second-order mean balance, expand the velocity and friction:
Averaging the exact pressure derivative gives zero, so . Hence the weak peristaltic pumping of a collapsible tube result is
The mean laboratory volumetric flow rate is , forward with the wave when . The negative leading relative flux is therefore consistent with positive mean pumping.