Solution

ID: past-exam-of-the-mathematics-and-the-environment-and-mathematical-physiology-course-of-the-university-of-oxford/2009/b/6/d/iii/solution

For real , draw and . The first is nonnegative and increasing, while the second is negative and approaches zero from below. They cannot intersect on the nonnegative real axis. At zero the characteristic function equals , so a stationary zero-root crossing is excluded as well.
Consequently a loss of stability from a stable regime cannot occur by a real eigenvalue crossing through zero. It must involve a conjugate pair crossing the imaginary axis at nonzero frequency, yielding oscillatory instability. This does not mean that every characteristic root is nonreal: negative real roots may exist and are harmless for stability. The graph and imaginary-axis conditions are shown together below.
Figure 1.
Respiratory delay feedback: no growing real root and the exact first oscillatory threshold
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