Solution

ID: past-exam-of-the-mathematics-course-of-the-university-of-cambridge/2004/iii/paper-39/1/ii/solution

Apply the law of total variance to each epoch. The Poisson distribution has equal conditional mean and variance, while . Hence
The independent mutation counts in coalescent epochs permit variance additivity for independent random variables. Thus
The first term is conditional Poisson distribution variation; the second comes from the random lengths of the ancestral tree. These are the moments of the mutation count on a neutral coalescent tree.

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