An age-structured population model tracks a density transported toward greater age, reduced by age-dependent mortality, and replenished at age zero by births.
For birth rate , death rate and exponential population growth rate , the Euler-Lotka equation is
For annual survival probabilities , age-specific offspring numbers , and a population multiplier , the discrete Euler-Lotka equation is
Its value at is the expected lifetime offspring count of a newborn under the fixed vital rates. A value above one implies a growing mode , while a value below one implies decline.

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