Reciprocal heritable evolutionary responses in interacting lineages, each altering selection on the other. Functional matching, co-occurrence and immediate plastic responses are insufficient by themselves to demonstrate coevolution.
Reciprocal evolutionary change driven by opposed fitness effects, including some host–parasite and sexual interactions. Correlated traits or temporal change alone do not establish reciprocal selection.

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Coevolution refers to the process in which two or more species influence each other's evolutionary trajectory through selective pressures. This occurs when the interactions between these species lead to adaptations that benefit one or both parties, which can drive changes in their respective traits over time.