The columella is the central, column-like region of the root cap. In Arabidopsis thaliana, its cells are organized into recognizable files and tiers, distinct from the surrounding lateral root cap. Starch-containing amyloplasts in these cells can change position after the root is reoriented relative to gravity.[ref]
A 1998 study paired two kinds of evidence. Live imaging tracked amyloplast displacement, while laser ablation removed selected cap cells before measuring gravitropism. Inner central columella cells made a particularly strong contribution to the response and showed rapid amyloplast sedimentation. This agreement linked cellular geometry and a candidate sensing mechanism to whole-root behavior.[ref]
A settling particle alone is not a complete proof of sensing: movement might accompany some other process. Selective removal asks whether the relevant cells are needed, while simultaneous growth measurements help exclude a simple failure of root extension. The columella supplies sensory-associated tissue; much of the resulting bend occurs farther back in the root elongation zone.
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