Reinhold Benesch and Ruth E. Benesch, Nature 221, 618–622, 15 February 1969. DOI 10.1038/221618a0; PMID 5774935. Publisher abstract: 2,3-diphosphoglycerate in red cells, lowered oxygen affinity through preferential binding to deoxyhaemoglobin and one phosphate-metabolite molecule per haemoglobin tetramer. Full author names follow the publisher byline; no expansion of Ruth's middle initial is inferred.

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