Fix the sense of rotation by taking the fluid velocity to be , with . Its vorticity is . In the convention where is the gyrotactic reorientation time appearing with aligning rate , the bottom-heavy spherical-cell orientation dynamics is
For the stable orientation in the transverse plane, write . The angle equation is . If , its stable fixed point satisfies
The other planar fixed point has negative and is unstable. The positive- orientation also damps a small out-of-plane component because .
Once the orientation has relaxed to , the transverse trajectory obeys
Define the shifted axis by
Then rotates rigidly with angular speed , so its squared length is constant. These are the gyrotactic circular paths in a rotating cylinder, and
Here is the acute angle between the plane through the two parallel axes and the horizontal plane; the directed displacement in the chosen cross-section points to the lower-left. Reversing the sense of rotation reverses the horizontal displacement. Arbitrary initial orientations have an orientation transient before the exact circular trajectories; neglecting randomness alone does not remove that transient. The bulk result also assumes that the trajectory does not intersect the cylinder wall.

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