In the small-slope Monge representation, , , and the leading transverse velocity is . With no imposed axial prestress, the induced filament tension is second order in transverse amplitude, so its transverse contribution is higher order. The transverse component of force balance reduces to the small-slope elastohydrodynamic filament equationAn externally imposed zeroth-order tension would instead add .
For the transverse motion, the off-diagonal part of the resistive-force theory tensor gives axial propulsive force density to second order. Its integral isSubstitute the bending equation and integrate by parts:The boundary expression for transverse filament thrust is thereforeOnly endpoint slope, bending moment, and shear enter this expression. Exact inextensibility also generates second-order longitudinal material motion; if that motion is retained in instantaneous total axial drag, it adds . For a periodic deformation with fixed axial base position, that additional term has zero cycle average. Thus the displayed formula is the transverse propulsive contribution and also gives the cycle-averaged propulsive force.
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