At a nonzero fixed point, both bracketed per-capita growth rates vanish:
Hence the coexistence equilibrium of this Lotka-Volterra predator-prey model is
Since , division of each population equation by gives
Define the Hamiltonian function
and the antisymmetric matrix
Then
Along a solution, the chain rule and antisymmetry of give
because for every real vector . Therefore
The closed level curves around the strict minimum at the coexistence equilibrium are the periodic population cycles of the Lotka-Volterra predator-prey model.

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