At a nonzero fixed point, both bracketed per-capita growth rates vanish:Hence the coexistence equilibrium of this Lotka-Volterra predator-prey model is
Since , division of each population equation by givesDefine the Hamiltonian functionand the antisymmetric matrixThen
Along a solution, the chain rule and antisymmetry of givebecause for every real vector . ThereforeThe closed level curves around the strict minimum at the coexistence equilibrium are the periodic population cycles of the Lotka-Volterra predator-prey model.
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