The cosmological continuity equation expresses conservation of dark-matter mass: is the density contrast, is the peculiar velocity, and a prime denotes a conformal time derivative. The cosmological Euler equation expresses momentum conservation: is the conformal Hubble rate, is the peculiar gravitational potential, is the density, and is the velocity-dispersion tensor of collisionless matter. The terms are respectively Hubble drag, convective acceleration, gravity, and velocity-dispersion stress.
For curl-free flow, introduce the peculiar-velocity divergence and set . A Fourier transform of the nonlinear continuity term gives
with the alpha mode-coupling kernel
Taking the divergence of the Euler equation gives the quadratic velocity kernel
This is the beta mode-coupling kernel; its symmetry follows from the two velocity factors.

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