Event-triggered harvesting by conserved energy ($\Delta H=-ch-d\log (1-h/X)$)

The predator-prey first integral $H=cX-d\log X+bY-a\log Y$ changes by the displayed amount when $h$ prey are removed. At a fixed prey threshold this increment is constant; sufficiently high thresholds reduce oscillation energy until the threshold is no longer crossed. At a fixed predator threshold, the increment depends on the prey abundance and on the crossing direction. A rising-predator balanced-quota cycle has negative return multiplier of magnitude greater than one; its leading small-oscillation approximation is marginal rather than a robust attractor.