The perturbations are density contrasts, and , evaluated in the common comoving synchronous gauge. For nonrelativistic cold dark matter of fixed particle mass, , hence . Thermal photons have photon number density and energy density . A small local temperature perturbation therefore gives
For adiabatic initial conditions, the ratio is unperturbed, so . Combining these equations proves
Equivalently , with and . The relation applies before causal pressure gradients can separate the two species on superhorizon scales; it need not remain true after cosmological horizon crossing.

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