Use the complete system in the PDF: the TeX omits . Both coordinate axes are invariant, and the equilibrium points are , and . The Jacobian matrix is
At its eigenvalues are , and at they are : both points are saddle equilibria. At the coexistence point,
Thus the interior point is a stable spiral. The nontrivial nullclines are and : increases below the first and decreases above it, while increases to the right of the second and decreases to the left. The rotation near coexistence is counterclockwise.
Figure 1.
Trajectories of a predator-prey system
.
A local linearization by itself does not prove that every interior solution has the same limit. For this logistic predator-prey model, let and use the Lyapunov function
The inequality makes nonnegative, with equality only at coexistence. Its sublevel sets are compact inside the positive quadrant, because diverges both at the axes and at infinity. Direct differentiation cancels the predator-prey cross terms and gives
On one has , and remaining there requires . The largest invariant subset is therefore the coexistence point. The LaSalle invariance principle proves
On , decays as toward the origin. On , the logistic differential equation takes every to one. The origin itself stays fixed. These boundary trajectories are the exceptions to the interior convergence statement.
For , differentiate and use the angular derivative . The polar coordinates satisfy
Hence the rotation is clockwise. The origin is an unstable equilibrium, while is a limit cycle of period . Writing gives the logistic differential equation . For ,
All nonzero solutions tend to the unit-circle attracting limit cycle as , spiralling outward for and inward for . For the solution exists for every real time and approaches the origin as ; for it remains periodic in both time directions. For , the denominator first vanishes at
This finite-time blow-up of an ordinary differential equation occurs backward in time. Such exterior solutions have no behavior: their maximal time interval is . The zero solution exists for all time and has no defined polar angle.

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