Up to a parameter-independent multinomial factor, the likelihood function is . A Lagrange multiplier for gives , whence and
Since all counts are positive, the log-likelihood function is strictly concave on the positive simplex and tends to minus infinity at its boundary, proving this is the unique maximum.
Under independence of random variables, , so the log-likelihood function separates as . Applying the same constrained maximization twice gives , , and
Twice the difference between the unrestricted and null maximized log-likelihood functions is
the required likelihood-ratio test statistic. Under the null hypothesis, its asymptotic chi-squared distribution has statistical degrees of freedom, the parameter-count difference . Here that is four. Since , reject independence at the 0.1 percent significance level. For four statistical degrees of freedom, the asymptotic upper-tail probability is . This establishes an association in these data; it is not a causal conclusion.
Write for individual survival probability. Conditional on a clump of size , its death probability is . Averaging with the geometric distribution gives
Thus
Here and ; different clumps and experimental repetitions are understood independent for the sampling model.
The live-clump counts have independent binomial distributions, all with the same . Up to a constant, their combined likelihood function is , maximized at when . Solving for gives . Therefore
Monotonicity of in makes this the maximum-likelihood estimator. At the sampling boundaries, gives , while gives only a supremum as , with no finite maximizer. These are the limiting interpretations of the formula.

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